Commitments and Contingencies - Contingencies - Additional Information (Details) $ in Thousands		1 Months Ended
	Dec. 01, 2020 claim	Dec. 31, 2020 USD ($) shares	Sep. 30, 2020 USD ($)	Sep. 30, 2021 USD ($) patent	Nov. 30, 2020 USD ($) claim
Loss Contingencies [Line Items]
Deferred revenue		$ 25,823		$ 26,716
Collaboration and licensing agreements
Loss Contingencies [Line Items]
Deferred revenue		$ 23,420		$ 23,040
Exotech, Inc.; AD Skincare, Inc.; and Thrive Agrobiotics, Inc.
Loss Contingencies [Line Items]
Investment					$ 326
XY, LLC | Licensing and patent infringement suit
Loss Contingencies [Line Items]
Loss contingency, claims settled, number | claim	1
Claims pending resolution | claim					2
Loss contingency, number of patents | patent				5
Division Of Enforcement Of The Securities And Exchange Commission | Methane Bioconversion Platform Disclosures
Loss Contingencies [Line Items]
Payments for civil money penalty			$ 2,500
Harvest Intrexon Enterprise Fund I, LP
Loss Contingencies [Line Items]
Stock issued in conjunction with settlement agreement, shares | shares		2,117,264
Shares issued in conjunction with settlement agreement		$ 18,103
Exotech, Inc.; AD Skincare, Inc.; and Thrive Agrobiotics, Inc. | Collaboration and licensing agreements
Loss Contingencies [Line Items]
Deferred revenue					$ 6,993