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Property, Plant and Equipment, Net
9 Months Ended
Sep. 30, 2020
Property, Plant and Equipment [Abstract]  
Property, Plant and Equipment, Net Property, Plant and Equipment, Net
Property, plant and equipment consist of the following:
September 30,
2020
December 31,
2019
Land and land improvements$9,845 $9,814 
Buildings and building improvements11,765 11,765 
Furniture and fixtures1,380 1,315 
Equipment48,028 54,448 
Leasehold improvements8,785 12,821 
Breeding stock5,533 5,191 
Computer hardware and software9,040 9,434 
Construction and other assets in progress3,670 5,313 
98,046 110,101 
Less: Accumulated depreciation and amortization(53,361)(49,132)
Property, plant and equipment, net$44,685 $60,969 
Depreciation expense was $2,195 and $2,874 for the three months ended September 30, 2020 and 2019, respectively, and $8,026 and $8,984 for the nine months ended September 30, 2020 and 2019, respectively.
During the second quarter of 2020, the Company suspended MBP Titan's operations. As a result, the Company reviewed the related property, plant and equipment and right-of-use assets for impairment. Based on the estimated undiscounted cash flows, the Company determined that the related asset values were not fully recoverable and calculated estimated fair values using market participant assumptions and discounted cash flow models. The estimated fair values were lower than the carrying values, and the Company recorded impairment losses of $9,914 related to property, plant, and equipment and $2,492 related to the right-of-use assets, which are included in impairment of other noncurrent assets on the accompanying condensed consolidated statement of operations for the nine months ended September 30, 2020.
During the third quarter of 2020, the Company recorded impairment losses of $920, which is included in impairment of other noncurrent assets on the accompanying condensed consolidated statements of operations for the three and nine months ended September 30, 2020, primarily related to right-of-use assets at certain of the Company's leased locations.
During the three and nine months ended September 30, 2019, the Company recorded $448 of property, plant and equipment impairment losses in conjunction with the closing of two of its reporting units during the third quarter of 2019, which is included in impairment of other noncurrent assets on the accompanying condensed consolidated statements of operations.