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Property, Plant and Equipment, Net
 6 Months Ended
Jun. 30, 2020
Property, Plant and Equipment [Abstract]  
Property, Plant and Equipment, Net Property, Plant and Equipment, Net
Property, plant and equipment consist of the following:
June 30,
2020		December 31,
2019
Land and land improvements$9,814  $9,814  
Buildings and building improvements11,765  11,765  
Furniture and fixtures1,293  1,315  
Equipment47,456  54,448  
Leasehold improvements9,016  12,821  
Breeding stock5,587  5,191  
Computer hardware and software8,922  9,434  
Construction and other assets in progress4,799  5,313  
98,652  110,101  
Less: Accumulated depreciation and amortization(51,696) (49,132) 
Property, plant and equipment, net$46,956  $60,969  
Depreciation expense was $2,900 and $2,932 for the three months ended June 30, 2020 and 2019, respectively, and $5,831 and $6,110 for the six months ended June 30, 2020 and 2019, respectively.
During the three months ended June 30, 2020, the Company suspended MBP Titan's operations. As a result, the Company reviewed the related property, plant and equipment and right-of-use assets for impairment. Based on the estimated undiscounted cash flows, the Company determined that the related asset values were not fully recoverable and calculated estimated fair values using market participant assumptions and discounted cash flow models. The estimated fair values were lower than the carrying values, and the Company recorded impairment losses of $9,914 related to property, plant, and equipment and $2,492 related to the right-of-use assets, which are included in impairment of assets on the accompanying condensed consolidated statements of operations.