Collaboration and Licensing Revenue - Summary of Deferred Revenue by Collaborator, Current Year (Details) - USD ($) $ in Thousands		Mar. 31, 2020	Dec. 31, 2019
Upfront and milestone payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue			$ 50,593
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2020-04-01 | Upfront and milestone payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue		$ 40,209
Oragenics, Inc. | Upfront and milestone payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue			2,864
Oragenics, Inc. | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2020-04-01
Contract With Customer, Asset And Liability [Line Items]
Average remaining performance period (in years)		4 years 2 months 12 days
Oragenics, Inc. | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2020-04-01 | Upfront and milestone payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue		$ 2,823
Intrexon Energy Partners, LLC | Upfront and milestone payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue			8,362
Intrexon Energy Partners, LLC | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2020-04-01
Contract With Customer, Asset And Liability [Line Items]
Average remaining performance period (in years)		4 years
Intrexon Energy Partners, LLC | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2020-04-01 | Upfront and milestone payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue		$ 8,362
Intrexon Energy Partners II, LLC | Upfront and milestone payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue			12,843
Intrexon Energy Partners II, LLC | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2020-04-01
Contract With Customer, Asset And Liability [Line Items]
Average remaining performance period (in years)		4 years 8 months 12 days
Intrexon Energy Partners II, LLC | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2020-04-01 | Upfront and milestone payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue		$ 12,843
Fibrocell Science, Inc. | Upfront and milestone payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue			17,697
Fibrocell Science, Inc. | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2020-04-01
Contract With Customer, Asset And Liability [Line Items]
Average remaining performance period (in years)		1 year
Fibrocell Science, Inc. | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2020-04-01 | Upfront and milestone payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue		$ 7,359
Harvest start-up entities | Upfront and milestone payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue	[1]		6,993
Harvest start-up entities | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2020-04-01
Contract With Customer, Asset And Liability [Line Items]
Average remaining performance period (in years)	[1]	4 years 10 months 24 days
Harvest start-up entities | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2020-04-01 | Upfront and milestone payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue	[1]	$ 6,993
Other | Upfront and milestone payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue			$ 1,834
Other | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2020-04-01
Contract With Customer, Asset And Liability [Line Items]
Average remaining performance period (in years)		2 years 6 months
Other | Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2020-04-01 | Upfront and milestone payments
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue		$ 1,829

[1] As of March 31, 2020 and December 31, 2019, the balance of deferred revenue for collaborations with Harvest start-up entities includes: Thrive Agrobiotics, Inc.; Exotech Bio, Inc.; and AD Skincare, Inc.