Collaboration and Licensing Revenue - Summary of Deferred Revenue (Details) - USD ($) $ in Thousands	Mar. 31, 2020	Dec. 31, 2019
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue	$ 44,018	$ 53,833
Current portion of deferred revenue	11,141	5,697
Long-term portion of deferred revenue	32,877	48,136
Collaboration and licensing agreements
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue	40,209	50,593
Prepaid product and service revenues
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue	3,461	2,805
Other
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue	$ 348	$ 435