Property, Plant and Equipment, Net - Schedule of Property, Plant and Equipment (Details) - USD ($) $ in Thousands	Sep. 30, 2019	Dec. 31, 2018
Property, Plant and Equipment [Abstract]
Land and land improvements	$ 11,011	$ 12,490
Buildings and building improvements	11,607	20,371
Furniture and fixtures	1,388	1,891
Equipment	67,020	74,555
Leasehold improvements	30,722	28,289
Breeding stock	4,867	4,582
Computer hardware and software	11,385	11,697
Trees	16,728	11,910
Construction and other assets in progress	26,319	18,880
Property, plant and equipment, gross	181,047	184,665
Less: Accumulated depreciation and amortization	(58,341)	(55,791)
Property, plant and equipment, net	$ 122,706	$ 128,874

Amount of accumulated depreciation, depletion and amortization for physical assets used in the normal conduct of business to produce goods and services.

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Amount before accumulated depreciation of building structures held for productive use including addition, improvement, or renovation to the structure, including, but not limited to, interior masonry, interior flooring, electrical, and plumbing.

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Amount of structure or a modification to a structure under construction. Includes recently completed structures or modifications to structures that have not been placed into service.

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Amount before accumulated depreciation of equipment commonly used in offices and stores that have no permanent connection to the structure of a building or utilities. Examples include, but are not limited to, desks, chairs, tables, and bookcases.

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Amount before accumulated deprecation and depletion of real estate held for productive use and additions or improvements to real estate held for productive use, examples include, but are not limited to, walkways, driveways, fences, and parking lots. Excludes land held for sale.

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Amount before accumulated depreciation of additions or improvements to assets held under a lease arrangement.

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Amount before accumulated depreciation of tangible personal property used to produce goods and services, including, but is not limited to, tools, dies and molds, computer and office equipment.

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Amount before accumulated depreciation, depletion and amortization of physical assets used in the normal conduct of business and not intended for resale. Examples include, but are not limited to, land, buildings, machinery and equipment, office equipment, and furniture and fixtures.

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Amount before accumulated depreciation of animals maintained to create offspring or to produce goods to be sold for profit.

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Amount before accumulated depreciation and amortization of computer hardware and purchased software.

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Amount before accumulated depreciation of producing trees and trees in process.

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