Collaboration and Licensing Revenue - Summary of Deferred Revenue (Details) - USD ($) $ in Thousands	Sep. 30, 2019	Dec. 31, 2018
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue	$ 79,124	$ 69,764
Current portion of deferred revenue	12,764	15,554
Long-term portion of deferred revenue	66,360	54,210
Collaboration and licensing agreements
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue	74,431	63,284
Prepaid product and service revenues
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue	3,054	2,933
Other
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue	$ 1,639	$ 3,547