Collaboration and Licensing Revenue - Surterra Collaboration (Details) - Surterra Holdings, Inc. $ in Thousands	1 Months Ended
	Jun. 30, 2019 USD ($)
Collaboration and licensing agreements
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Collaborative arrangement consideration received, cash	$ 10,000
Collaborative arrangement consideration received, value	$ 4,530
Collaboration agreement termination notice period	90 days
Maximum
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Milestone payments required upon successful achievement	$ 68,000