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Summary of Significant Accounting Policies - Equity Method Investments - Additional Information (Details) - USD ($)
$ in Thousands
3 Months Ended 9 Months Ended
Sep. 30, 2019
Sep. 30, 2018
Sep. 30, 2019
Sep. 30, 2018
AquaBounty Technologies, Inc.        
Schedule of Equity Method Investments [Line Items]        
Equity method investments, fair value disclosure $ 16,320   $ 16,320  
Ownership interest 38.00%   38.00%  
Unrealized appreciation (depreciation) in fair value of equity securities $ (3,721)   $ 2,081  
Oragenics, Inc.        
Schedule of Equity Method Investments [Line Items]        
Unrealized appreciation (depreciation) in fair value of equity securities   $ (387)   $ (1,547)