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Mergers and Acquisitions - Asset Acquisition of Certain Harvest Entities - Additional Information (Details) - USD ($)
1 Months Ended
Sep. 30, 2018
Jun. 30, 2019
Dec. 31, 2018
Aug. 31, 2018
Business Acquisition [Line Items]        
Investments in affiliates   $ 18,093,000 $ 18,859,000  
Deferred revenue   $ 83,135,000 $ 69,764,000  
Harvest Intrexon Enterprise Fund I, LP        
Business Acquisition [Line Items]        
Cash received in asset acquisition $ 15,500,000      
Harvest Intrexon Enterprise Fund I, LP | ActoBio Therapeutics Inc.        
Business Acquisition [Line Items]        
Long-term debt issued to a related party in an asset acquisition 30,000,000      
CRS Bio, Inc.; Genten Therapeutics, Inc.; and Relive Genetics, Inc.        
Business Acquisition [Line Items]        
Investments in affiliates       $ 4,303,000
Ownership interest   100.00%    
Deferred revenue 10,078,000      
Write-off of in-process research and development acquired in asset acquisition $ 8,721,000