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Property, Plant and Equipment, Net
6 Months Ended
Jun. 30, 2019
Property, Plant and Equipment [Abstract]  
Property, Plant and Equipment, Net Property, Plant and Equipment, Net
Property, plant and equipment consist of the following:
 
June 30,
2019
 
December 31,
2018
Land and land improvements
$
11,674

 
$
12,490

Buildings and building improvements
11,590

 
20,371

Furniture and fixtures
1,684

 
1,891

Equipment
68,138

 
74,555

Leasehold improvements
31,925

 
28,289

Breeding stock
4,675

 
4,582

Computer hardware and software
12,131

 
11,697

Trees
16,141

 
11,910

Construction and other assets in progress
23,000

 
18,880

 
180,958

 
184,665

Less: Accumulated depreciation and amortization
(60,557
)
 
(55,791
)
Property, plant and equipment, net
$
120,401

 
$
128,874


The deconsolidation of AquaBounty (Note 1) in April 2019 resulted in the reduction of $24,186 of property, plant and equipment, net on the accompanying consolidated balance sheet as of June 30, 2019.
During the three and six months ended June 30, 2018, the Company recorded a $4,972 loss on disposal of certain leasehold improvements, equipment, and other fixed assets, in conjunction with the closing of one of its research and development facilities in Brazil.
Depreciation expense was $3,347 and $3,642 for the three months ended June 30, 2019 and 2018, respectively, and $6,920 and $7,098 for the six months ended June 30, 2019 and 2018, respectively.