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Property, Plant and Equipment, Net
3 Months Ended
Mar. 31, 2018
Property, Plant and Equipment [Abstract]  
Property, Plant and Equipment, Net
Property, Plant and Equipment, Net
Property, plant and equipment consist of the following:
 
March 31,
2018
 
December 31,
2017
Land and land improvements
$
11,815

 
$
11,767

Buildings and building improvements
18,160

 
18,183

Furniture and fixtures
2,516

 
2,515

Equipment
69,568

 
65,863

Leasehold improvements
27,811

 
25,277

Breeding stock
4,204

 
3,832

Computer hardware and software
10,442

 
10,128

Trees
7,996

 
6,642

Construction and other assets in progress
15,819

 
14,113

 
168,331

 
158,320

Less: Accumulated depreciation and amortization
(49,087
)
 
(45,646
)
Property, plant and equipment, net
$
119,244

 
$
112,674


Depreciation expense was $3,456 and $2,782 for the three months ended March 31, 2018 and 2017, respectively.