Property, Plant and Equipment, Net - Schedule of Property, Plant and Equipment (Details) - USD ($) $ in Thousands	Dec. 31, 2017	Dec. 31, 2016
Property, Plant and Equipment [Abstract]
Land and land improvements	$ 11,767	$ 10,904
Buildings and building improvements	18,183	8,123
Furniture and fixtures	2,515	2,176
Equipment	65,863	44,392
Leasehold improvements	25,277	15,105
Breeding stock	3,832	3,893
Computer hardware and software	10,128	6,844
Trees	6,642	2,772
Construction and other assets in progress	14,113	4,513
Property, plant and equipment, gross	158,320	98,722
Less: Accumulated depreciation and amortization	(45,646)	(34,050)
Property, plant and equipment, net	$ 112,674	$ 64,672