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Property, Plant and Equipment, Net
 12 Months Ended
Dec. 31, 2017
Property, Plant and Equipment [Abstract]  
Property, Plant and Equipment, Net
Property, Plant and Equipment, Net
Property, plant and equipment consist of the following:
 
December 31,
 
2017
 
2016
Land and land improvements
$
11,767

 
$
10,904

Buildings and building improvements
18,183

 
8,123

Furniture and fixtures
2,515

 
2,176

Equipment
65,863

 
44,392

Leasehold improvements
25,277

 
15,105

Breeding stock
3,832

 
3,893

Computer hardware and software
10,128

 
6,844

Trees
6,642

 
2,772

Construction and other assets in progress
14,113

 
4,513

 
158,320

 
98,722

Less: Accumulated depreciation and amortization
(45,646
)
 
(34,050
)
Property, plant and equipment, net
$
112,674

 
$
64,672


Depreciation expense was $11,951, $9,387 and $7,872 for the years ended December 31, 2017, 2016 and 2015, respectively.
Included in the table above is $14,219 of land, buildings, and equipment related to a 2017 asset acquisition of a land-based aquaculture facility to be used in the production of AquAdvantage salmon in Indiana.