Property, Plant and Equipment, Net - Schedule of Property, Plant and Equipment (Details) - USD ($) $ in Thousands	Sep. 30, 2017	Dec. 31, 2016
Property, Plant and Equipment [Abstract]
Land and land improvements	$ 11,642	$ 10,904
Buildings and building improvements	15,216	8,123
Furniture and fixtures	2,220	2,176
Equipment	58,851	44,392
Leasehold improvements	22,974	15,105
Breeding stock	3,822	3,893
Computer hardware and software	9,817	6,844
Trees	5,719	2,772
Construction and other assets in progress	15,138	4,513
Property, plant and equipment, gross	145,399	98,722
Less: Accumulated depreciation and amortization	(42,523)	(34,050)
Property, plant and equipment, net	$ 102,876	$ 64,672