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Consolidated Statements of Operations (Parenthetical) - USD ($)
$ in Thousands
3 Months Ended 9 Months Ended
Sep. 30, 2017
Sep. 30, 2016
Sep. 30, 2017
Sep. 30, 2016
Related Party Transaction [Line Items]        
Collaboration and licensing revenues $ 28,155 $ 30,590 $ 89,384 $ 82,144
Related Parties, Aggregated        
Related Party Transaction [Line Items]        
Collaboration and licensing revenues $ 24,492 $ 26,688 $ 77,937 $ 70,299