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Goodwill and Intangible Assets, Net
9 Months Ended
Sep. 30, 2017
Goodwill and Intangible Assets Disclosure [Abstract]  
Goodwill and Intangible Assets, Net
Goodwill and Intangible Assets, Net
The changes in the carrying amount of goodwill for the nine months ended September 30, 2017 are as follows:
Balance at December 31, 2016
$
157,175

Acquisitions
4,850

Foreign currency translation adjustments
4,796

Balance at September 30, 2017
$
166,821


No accumulated impairment losses existed as of September 30, 2017 and December 31, 2016.
Intangible assets consist of the following as of September 30, 2017:
 
Weighted Average Useful Life (Years)
 
Gross Carrying Amount
 
Accumulated Amortization
 
Net
Patents, developed technologies and know-how
15.7
 
$
266,697

 
$
(43,465
)
 
$
223,232

Customer relationships
6.5
 
10,700

 
(6,033
)
 
4,667

Trademarks
9.3
 
6,800

 
(2,373
)
 
4,427

In-process research and development
 
 
8,571

 
—

 
8,571

Total
 
 
$
292,768

 
$
(51,871
)
 
$
240,897


Intangible assets consist of the following as of December 31, 2016:
 
Gross Carrying Amount
 
Accumulated Amortization
 
Net
Patents, developed technologies and know-how
$
236,401

 
$
(29,748
)
 
$
206,653

Customer relationships
10,700

 
(4,672
)
 
6,028

Trademarks
6,800

 
(1,792
)
 
5,008

Covenant not to compete
370

 
(339
)
 
31

In-process research and development
7,895

 
—

 
7,895

Total
$
262,166

 
$
(36,551
)
 
$
225,615


The balance of in-process research and development includes certain in-process research and development technology acquired in the Company's acquisition of Oxitec in September 2015, and amortization will begin once certain regulatory approvals have been obtained for the in-process programs.
Amortization expense was $5,001 and $3,651 for the three months ended September 30, 2017 and 2016, respectively, and $14,258 and $10,888 for the nine months ended September 30, 2017 and 2016, respectively.