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Consolidated Statements of Operations (Parenthetical) - USD ($)
$ in Thousands
3 Months Ended 6 Months Ended
Jun. 30, 2017
Jun. 30, 2016
Jun. 30, 2017
Jun. 30, 2016
Related Party Transaction [Line Items]        
Collaboration and licensing revenues $ 28,164 $ 27,481 $ 61,229 $ 51,554
Related Parties, Aggregated        
Related Party Transaction [Line Items]        
Collaboration and licensing revenues $ 24,558 $ 23,612 $ 53,445 $ 43,611