Related Party Transactions (Details) - USD ($) $ in Thousands	1 Months Ended	3 Months Ended
	Nov. 30, 2015	Mar. 31, 2017	Mar. 31, 2016	Dec. 31, 2016
Related Party Transaction [Line Items]
Shares issued as payment for services		$ 2,915
Fair value of financial assets measured at fair value on a recurring basis		293,831		$ 335,307
Other nonoperating income		595	$ 561
Third Security, LLC
Related Party Transaction [Line Items]
Expense for services		$ 66	$ 46
Initial term of services agreement	1 year
Shares issued as payment for services (in shares)		103,930	79,870
Shares issued as payment for services		$ 2,039	$ 2,267
Third Security, LLC | Common Stock
Related Party Transaction [Line Items]
Expense for services	$ 800
Harvest Intrexon Enterprise Fund I, LP
Related Party Transaction [Line Items]
Other nonoperating income		603	$ 645
Convertible Note and Warrants | Other Assets | Fibrocell Science, Inc.
Related Party Transaction [Line Items]
Fair value of financial assets measured at fair value on a recurring basis		$ 2,315		$ 1,642
Third Security, LLC | Chief Executive Officer
Related Party Transaction [Line Items]
Ownership interest		100.00%