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Goodwill and Intangible Assets, Net
3 Months Ended
Mar. 31, 2017
Goodwill and Intangible Assets Disclosure [Abstract]  
Goodwill and Intangible Assets, Net
Goodwill and Intangible Assets, Net
The changes in the carrying amount of goodwill for the three months ended March 31, 2017 are as follows:
Balance at December 31, 2016
$
157,175

Foreign currency translation adjustments
650

Balance at March 31, 2017
$
157,825


No goodwill or accumulated impairment losses existed as of March 31, 2017 and December 31, 2016.
Intangible assets consist of the following at March 31, 2017:
 
Weighted Average Useful Life (Years)
 
Gross Carrying Amount
 
Accumulated Amortization
 
Net
Patents, developed technologies and know-how
15.9
 
$
238,513

 
$
(33,789
)
 
$
204,724

Customer relationships
6.5
 
10,700

 
(5,156
)
 
5,544

Trademarks
9.3
 
6,800

 
(1,986
)
 
4,814

In-process research and development
 
 
7,992

 

 
7,992

Total
 
 
$
264,005

 
$
(40,931
)
 
$
223,074


Intangible assets consist of the following at December 31, 2016:
 
Gross Carrying Amount
 
Accumulated Amortization
 
Net
Patents, developed technologies and know-how
$
236,401

 
$
(29,748
)
 
$
206,653

Customer relationships
10,700

 
(4,672
)
 
6,028

Trademarks
6,800

 
(1,792
)
 
5,008

Covenant not to compete
370

 
(339
)
 
31

In-process research and development
7,895

 

 
7,895

Total
$
262,166

 
$
(36,551
)
 
$
225,615


The balance of in-process research and development as of March 31, 2017 includes certain in-process research and development technology acquired in the Company's acquisition of Oxitec in September 2015, and amortization will begin once certain regulatory approvals have been obtained for the in-process programs.
Amortization expense was $4,618 and $3,515 for the three months ended March 31, 2017 and 2016, respectively.