Mergers and Acquisitions - Trans Ova - Additional Information (Details) - USD ($)	1 Months Ended		3 Months Ended								12 Months Ended
	Feb. 28, 2015	Aug. 31, 2014	Dec. 31, 2016	Sep. 30, 2016	Jun. 30, 2016	Mar. 31, 2016	Dec. 31, 2015	Sep. 30, 2015	Jun. 30, 2015	Mar. 31, 2015	Dec. 31, 2016	Dec. 31, 2015	Dec. 31, 2014
Business Acquisition [Line Items]
Deferred cash consideration											$ 0	$ 1,992,000	$ 20,115,000
Revenues			$ 46,002,000	$ 48,985,000	$ 52,501,000	$ 43,438,000	$ 41,498,000	$ 53,367,000	$ 44,891,000	$ 33,849,000	190,926,000	173,605,000	71,930,000
Net income (loss)			(44,912,000)	$ (30,011,000)	$ (50,031,000)	$ (65,320,000)	$ (33,275,000)	$ (39,029,000)	$ (41,494,000)	$ 25,804,000	(190,274,000)	(87,994,000)	(85,616,000)
Trans Ova Genetics, LC
Business Acquisition [Line Items]
Percentage of outstanding common stock acquired		100.00%
Business combination, consideration paid, shares issued		1,444,388
Cash		$ 63,625,000
Deferred cash consideration		$ 20,115,000
Revenues											78,143,000	83,912,000	26,352,000
Net income (loss)											(9,679,000)	$ (1,531,000)	2,000
Future contingent payments based on revenue targets			3,000,000								3,000,000
Current contingent consideration liability			$ 0								$ 0
Exemplar Genetics, LLC
Business Acquisition [Line Items]
Cash paid to acquire minority interest	$ 1,566,000
Shares issued to acquire minority interest	307,074
Minimum
Business Acquisition [Line Items]
Expected useful life of intangible asset											2 years
Minimum | Trans Ova Genetics, LC
Business Acquisition [Line Items]
Expected useful life of intangible asset		3 years
Maximum
Business Acquisition [Line Items]
Expected useful life of intangible asset											21 years
Maximum | Trans Ova Genetics, LC
Business Acquisition [Line Items]
Expected useful life of intangible asset		9 years
Selling, general and administrative | Trans Ova Genetics, LC
Business Acquisition [Line Items]
Business combination, acquisition related cost													$ 713,000