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Mergers and Acquisitions - Trans Ova - Fair Value of Consideration Transferred (Details) - USD ($)
$ in Thousands
1 Months Ended 12 Months Ended
Aug. 31, 2014
Dec. 31, 2016
Dec. 31, 2015
Dec. 31, 2014
Business Acquisition [Line Items]        
Deferred cash consideration   $ 0 $ 1,992 $ 20,115
Trans Ova Genetics, LC        
Business Acquisition [Line Items]        
Cash $ 63,625      
Common shares 32,802      
Deferred cash consideration 20,115      
Total consideration transferred 116,542      
Fair value of noncontrolling interest 11,333      
Total $ 127,875