Mergers and Acquisitions - Fair Value of Assets Acquired and Liabilities Assumed (Details) - USD ($) $ in Thousands	Sep. 30, 2016	Dec. 31, 2015	Sep. 30, 2015	Apr. 30, 2015	Feb. 28, 2015
Business Acquisition [Line Items]
Goodwill	$ 159,793	$ 165,169
Oxitec
Business Acquisition [Line Items]
Cash			$ 3,780
Trade receivables			125
Other receivables			7,395
Prepaid expenses and other			121
Property, plant and equipment			1,198
Intangible assets			96,854
Total assets acquired			109,473
Accounts payable			1,187
Accrued compensation and benefits			246
Other accrued liabilities			210
Deferred revenue			120
Deferred tax liabilities			12,584
Total liabilities assumed			14,347
Net assets acquired			95,126
Goodwill			51,268
Total consideration			$ 146,394
Okanagan
Business Acquisition [Line Items]
Cash				$ 58
Trade receivables				16
Other receivables				49
Property, plant and equipment				32
Intangible assets				36,500
Total assets acquired				36,655
Accounts payable				181
Deferred revenue				181
Deferred tax liabilities				8,847
Total liabilities assumed				9,209
Net assets acquired				27,446
Goodwill				13,487
Total consideration				$ 40,933
ActoGeniX NV
Business Acquisition [Line Items]
Cash					$ 3,180
Other receivables					305
Prepaid expenses and other					31
Property, plant and equipment					209
Intangible assets					68,100
Other non-current assets					23
Total assets acquired					71,848
Accounts payable					230
Accrued compensation and benefits					196
Other accrued liabilities					253
Deferred revenue					732
Deferred tax liabilities					612
Total liabilities assumed					2,023
Net assets acquired					69,825
Goodwill					2,649
Total consideration					$ 72,474