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Collaboration and Licensing Revenue - Summarized Collaboration and Licensing Revenues (Details) - USD ($)
$ in Thousands
3 Months Ended 6 Months Ended
Jun. 30, 2016
Jun. 30, 2015
Jun. 30, 2016
Jun. 30, 2015
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues $ 27,481 $ 17,181 $ 51,554 $ 31,964
Upfront and Milestone Payments        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 8,055 3,703 13,827 6,734
Research and Development Services        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 19,426 13,478 37,727 25,230
ZIOPHARM Oncology, Inc.        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 6,970 5,250 13,951 9,051
ZIOPHARM Oncology, Inc. | Upfront and Milestone Payments        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 922 644 1,844 1,288
ZIOPHARM Oncology, Inc. | Research and Development Services        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 6,048 4,606 12,107 7,763
Oragenics, Inc.        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 507 375 1,313 645
Oragenics, Inc. | Upfront and Milestone Payments        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 261 307 524 569
Oragenics, Inc. | Research and Development Services        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 246 68 789 76
Fibrocell Science, Inc.        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 1,394 1,918 3,251 4,079
Fibrocell Science, Inc. | Upfront and Milestone Payments        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 605 448 1,210 896
Fibrocell Science, Inc. | Research and Development Services        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 789 1,470 2,041 3,183
Genopaver, LLC        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 1,637 935 3,215 1,604
Genopaver, LLC | Upfront and Milestone Payments        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 68 68 137 137
Genopaver, LLC | Research and Development Services        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 1,569 867 3,078 1,467
S & I Ophthalmic, LLC        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 2,358 890 3,544 1,645
S & I Ophthalmic, LLC | Upfront and Milestone Payments        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 0 0 0 0
S & I Ophthalmic, LLC | Research and Development Services        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 2,358 890 3,544 1,645
OvaXon, LLC        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 808 662 1,502 1,306
OvaXon, LLC | Upfront and Milestone Payments        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 0 0 0 0
OvaXon, LLC | Research and Development Services        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 808 662 1,502 1,306
Intrexon Energy Partners, LLC        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 4,212 3,356 8,200 6,166
Intrexon Energy Partners, LLC | Upfront and Milestone Payments        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 625 625 1,250 1,250
Intrexon Energy Partners, LLC | Research and Development Services        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 3,587 2,731 6,950 4,916
Persea Bio, LLC        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 331 266 655 506
Persea Bio, LLC | Upfront and Milestone Payments        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 125 125 250 250
Persea Bio, LLC | Research and Development Services        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 206 141 405 256
Ares Trading S.A.        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 2,218 739 4,623 739
Ares Trading S.A. | Upfront and Milestone Payments        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 1,597 739 3,194 739
Ares Trading S.A. | Research and Development Services        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 621 0 1,429 0
Thrive Agrobiotics, Inc.        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 450   884  
Thrive Agrobiotics, Inc. | Upfront and Milestone Payments        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 46   92  
Thrive Agrobiotics, Inc. | Research and Development Services        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 404   792  
Intrexon Energy Partners II, LLC        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 894   1,444  
Intrexon Energy Partners II, LLC | Upfront and Milestone Payments        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 500   1,000  
Intrexon Energy Partners II, LLC | Research and Development Services        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 394   444  
Exotech Bio, Inc.        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 139   139  
Exotech Bio, Inc. | Upfront and Milestone Payments        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 139   139  
Exotech Bio, Inc. | Research and Development Services        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 0   0  
Relieve Genetics, Inc.        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 350   350  
Relieve Genetics, Inc. | Upfront and Milestone Payments        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 120   120  
Relieve Genetics, Inc. | Research and Development Services        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 230   230  
Intrexon T1D Partners, LLC        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 310   310  
Intrexon T1D Partners, LLC | Upfront and Milestone Payments        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 278   278  
Intrexon T1D Partners, LLC | Research and Development Services        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 32   32  
Other        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 4,903 2,790 8,173 6,223
Other | Upfront and Milestone Payments        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues 2,769 747 3,789 1,605
Other | Research and Development Services        
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]        
Collaboration and licensing revenues $ 2,134 $ 2,043 $ 4,384 $ 4,618