Mergers and Acquisitions - Oxitec - Additional Information (Details) - Oxitec Limited $ in Thousands	1 Months Ended
	Sep. 30, 2015 USD ($) shares
Business Acquisition [Line Items]
Percentage of outstanding common stock acquired	100.00%
Consideration transferred, shares issued or issuable | shares	1,359,343
Cash	$ 90,199
Consideration transferred, cash and working capital adjustments	$ 9,449
Equity interest issued or issuable, number of shares withheld as escrow | shares	480,422
Cash consideration withheld as escrow	$ 1,991
Consideration withheld as escrow, transfer period after closing	18 months
Consideration transferred	$ 146,394
Selling, general and administrative
Business Acquisition [Line Items]
Acquisition related costs	$ 1,675