Collaboration and Licensing Revenue - Additional Information (Details)	1 Months Ended
	Mar. 31, 2016 USD ($)
Exotech Bio, Inc.
Collaboration and Licensing Agreements [Line Items]
Maximum milestone payments required upon successful achievement, per product	$ 52,500,000
Required notice period for voluntary termination of collaborative agreement	90 days
Exotech Bio, Inc. | Upfront | Upfront and Milestone Payments
Collaboration and Licensing Agreements [Line Items]
Collaborative arrangement consideration received, value	$ 5,000,000
Relieve Genetics, Inc.
Collaboration and Licensing Agreements [Line Items]
Maximum milestone payments required upon successful achievement, per product	$ 52,500,000
Required notice period for voluntary termination of collaborative agreement	90 days
Relieve Genetics, Inc. | Upfront | Upfront and Milestone Payments
Collaboration and Licensing Agreements [Line Items]
Collaborative arrangement consideration received, value	$ 4,333,000
Intrexon T1D Partners, LLC
Collaboration and Licensing Agreements [Line Items]
Required notice period for voluntary termination of collaborative agreement	90 days
Intrexon T1D Partners, LLC | Upfront | Upfront and Milestone Payments
Collaboration and Licensing Agreements [Line Items]
Collaborative arrangement consideration received, value	$ 10,000,000