Collaboration and Licensing Revenue (Tables)

3 Months Ended

Mar. 31, 2016

Organization, Consolidation and Presentation of Financial Statements [Abstract]

Summarized Collaboration and Licensing Revenues

The following tables summarize the amounts recorded as revenue in the consolidated statements of operations for each significant collaboration or licensing agreement for the three months ended March 31, 2016 and 2015.

	Three Months Ended March 31, 2016
	Revenue Recognized From				Total
	Upfront and Milestone Payments		Research and Development Services
ZIOPHARM Oncology, Inc.	$	922	$	6,059	$	6,981
Oragenics, Inc.	263		543		806
Fibrocell Science, Inc.	605		1,252		1,857
Genopaver, LLC	69		1,509		1,578
S & I Ophthalmic, LLC	—		1,186		1,186
OvaXon, LLC	—		694		694
Intrexon Energy Partners, LLC	625		3,363		3,988
Persea Bio, LLC	125		199		324
Ares Trading S.A.	1,597		808		2,405
Thrive Agrobiotics, Inc.	46		388		434
Intrexon Energy Partners II, LLC	500		50		550
Other	1,020		2,250		3,270
Total	$	5,772	$	18,301	$	24,073

	Three Months Ended March 31, 2015
	Revenue Recognized From				Total
	Upfront and Milestone Payments		Research and Development Services
ZIOPHARM Oncology, Inc.	$	644	$	3,157	$	3,801
Oragenics, Inc.	262		8		270
Fibrocell Science, Inc.	448		1,713		2,161
Genopaver, LLC	69		600		669
S & I Ophthalmic, LLC	—		755		755
OvaXon, LLC	—		644		644
Intrexon Energy Partners, LLC	625		2,185		2,810
Persea Bio, LLC	125		115		240
Other	858		2,575		3,433
Total	$	3,031	$	11,752	$	14,783

Summary of Deferred Revenue

Deferred revenue consists of the following:

	March 31, 2016		December 31, 2015
Upfront and milestone payments	$	194,849	$	181,331
Prepaid research and development services	9,439		10,938
Prepaid product and service revenues	5,201		4,759
Other	749		701
Total	$	210,238	$	197,729
Current portion of deferred revenue	$	37,695	$	35,366
Long-term portion of deferred revenue	172,543		162,363
Total	$	210,238	$	197,729

Summary of Deferred Revenue by Collaborator

The following table summarizes the remaining balance of deferred revenue associated with upfront and milestone payments for each significant collaboration and licensing agreement:

	March 31, 2016		December 31, 2015
ZIOPHARM Oncology, Inc.	$	29,416	$	30,338
Oragenics, Inc.	8,550		8,813
Fibrocell Science, Inc.	20,840		21,445
Genopaver, LLC	2,181		2,250
Intrexon Energy Partners, LLC	20,000		20,625
Persea Bio, LLC	4,375		4,500
Ares Trading S.A.	51,970		53,567
Thrive Agrobiotics, Inc.	1,575		1,621
Intrexon Energy Partners II, LLC	17,333		17,833
Exotech Bio, Inc.	5,000		—
Relieve Genetics, Inc.	4,333		—
Intrexon T1D Partners, LLC	10,000		—
Other	19,276		20,339
Total	$	194,849	$	181,331