Mergers and Acquisitions - Additional Information (Details)	1 Months Ended						3 Months Ended		9 Months Ended
	Sep. 30, 2015 USD ($) shares	Aug. 31, 2015	Apr. 30, 2015 USD ($) shares	Feb. 28, 2015 USD ($) shares	Aug. 31, 2014 USD ($) installment shares	Mar. 31, 2014 USD ($) note shares	Sep. 30, 2015 USD ($)	Sep. 30, 2014 USD ($)	Sep. 30, 2015 USD ($)	Sep. 30, 2014 USD ($)	Sep. 30, 2012 USD ($)
Business Acquisition [Line Items]
Deferred cash consideration									$ 11,440,000	$ 20,115,000
Revenues							$ 53,367,000	$ 21,197,000	132,107,000	40,838,000
Net income (loss)							$ (39,029,000)	(53,862,000)	$ (54,719,000)	(103,548,000)
Oxitec Limited
Business Acquisition [Line Items]
Percentage of outstanding common stock acquired	100.00%						100.00%		100.00%
Consideration transferred, shares issued or issuable | shares	1,359,343
Cash	$ 90,199,000
Cash and working capital adjustments	$ 9,449,000
Payment of cash and working capital adjustments, number of days after closing	85 days
Consideration transferred, shares issuable withheld as escrow | shares	480,422
Consideration transferred, cash withheld as escrow	$ 1,991,000
Transfer of cash withheld as escrow, number of months after closing	18 months
Total consideration transferred	$ 146,394,000
Oxitec Limited | Selling, general and administrative
Business Acquisition [Line Items]
Business combination, acquisition related cost							$ 1,644,000		$ 1,644,000
Okanagan Specialty Fruits Inc
Business Acquisition [Line Items]
Percentage of outstanding common stock acquired			100.00%
Consideration transferred, shares issued or issuable | shares			707,853
Cash			$ 10,000,000
Total consideration transferred			$ 40,933,000
Business combination, acquisition related cost									341,000
Intangible assets, useful life			14 years
Okanagan Specialty Fruits Inc | Selling, general and administrative
Business Acquisition [Line Items]
Business combination, acquisition related cost									267,000
ActoGeniX NV
Business Acquisition [Line Items]
Percentage of outstanding common stock acquired				100.00%
Consideration transferred, shares issued or issuable | shares				965,377
Cash				$ 32,739,000
Total consideration transferred				72,474,000
Business combination, acquisition related cost									418,000
Intangible assets, useful life		18 years
ActoGeniX NV | Selling, general and administrative
Business Acquisition [Line Items]
Business combination, acquisition related cost									381,000
Trans Ova Genetics, LC
Business Acquisition [Line Items]
Percentage of outstanding common stock acquired					100.00%
Consideration transferred, shares issued or issuable | shares					1,444,388
Cash					$ 63,625,000
Total consideration transferred					116,542,000
Business combination, acquisition related cost										713,000
Deferred cash consideration					$ 20,115,000
Deferred cash consideration, number of payment installments | installment					3
Revenues							18,283,000	8,502,000	64,743,000	8,502,000
Net income (loss)							(3,655,000)	(481,000)	530,000	(481,000)
Total consideration transferred, including the noncontrolling interest					$ 127,875,000
Possible future contingent payments to former equity holders											$ 6,000,000
Possible future contingent payments to former equity holders, current valuation	$ 0						$ 0		$ 0
Trans Ova Genetics, LC | Selling, general and administrative
Business Acquisition [Line Items]
Business combination, acquisition related cost								$ 372,000		684,000
Exemplar Genetics, LLC
Business Acquisition [Line Items]
Cash paid to acquire noncontrolling interest				$ 1,566,000
Shares issued to acquire noncontrolling interest | shares				307,074
Medistem, Inc.
Business Acquisition [Line Items]
Percentage of outstanding common stock acquired						100.00%
Consideration transferred, shares issued or issuable | shares						714,144
Cash						$ 4,920,000
Total consideration transferred						$ 24,995,000
Business combination, acquisition related cost										680,000
Number of promissory notes settled upon the closing of the merger | note						2
Shares of merger consideration surrendered by acquiree management to reimburse for required payroll tax withholdings | shares						17,695
Medistem, Inc. | Settlement of Promissory Notes
Business Acquisition [Line Items]
Total consideration transferred						$ 707,000
Medistem, Inc. | Selling, general and administrative
Business Acquisition [Line Items]
Business combination, acquisition related cost										$ 310,000
Minimum | Trans Ova Genetics, LC
Business Acquisition [Line Items]
Intangible assets, useful life					3 years
Maximum | Trans Ova Genetics, LC
Business Acquisition [Line Items]
Intangible assets, useful life					9 years