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Mergers and Acquisitions - Fair Value of Consideration Transferred - Trans Ova (Details) (USD $) In Thousands, unless otherwise specified	12 Months Ended			0 Months Ended
	Dec. 31, 2014	Dec. 31, 2013	Dec. 31, 2012	Aug. 08, 2014
Business Acquisition [Line Items]
Deferred cash consideration	$ 20,115	$ 0	$ 0
Trans Ova Genetics, LC
Business Acquisition [Line Items]
Cash				63,625
Common shares				32,802
Deferred cash consideration				20,115
Total consideration transferred				116,542
Fair value of noncontrolling interest				11,333
Total				$ 127,875