Collaboration Revenue (Tables)

12 Months Ended

Dec. 31, 2014

Organization, Consolidation and Presentation of Financial Statements [Abstract]

Summarized Collaboration Revenues

The following table summarizes the amounts recorded in the consolidated statements of operations for each significant collaboration for the years ended December 31, 2014, 2013 and 2012.

	Year Ended December 31, 2014
	Collaboration Revenue Recognized From				Total
	Upfront and Milestone Payments		Research and Development Services
ZIOPHARM Oncology, Inc.	$	2,577	$	12,044	$	14,621
Synthetic Biologics, Inc.	651		273		924
Oragenics, Inc.	1,045		598		1,643
Fibrocell Science, Inc.	1,794		4,398		6,192
Genopaver, LLC	273		1,510		1,783
S & I Ophthalmic, LLC	—		2,832		2,832
OvaXon, LLC	—		2,799		2,799
Intrexon Energy Partners, LLC	1,875		4,227		6,102
Other	1,410		6,906		8,316
Total	$	9,625	$	35,587	$	45,212

	Year Ended December 31, 2013
	Collaboration Revenue Recognized From				Total
	Upfront and Milestone Payments		Research and Development Services
ZIOPHARM Oncology, Inc.	$	2,577	$	7,818	$	10,395
Synthetic Biologics, Inc.	2,187		1,048		3,235
Oragenics, Inc.	673		1,517		2,190
Fibrocell Science, Inc.	970		3,736		4,706
Genopaver, LLC	204		935		1,139
S & I Ophthalmic, LLC	—		417		417
Other	333		1,110		1,443
Total	$	6,944	$	16,581	$	23,525

	Year Ended December 31, 2012
	Collaboration Revenue Recognized From				Total
	Upfront and Milestone Payments		Research and Development Services
ZIOPHARM Oncology, Inc.	$	5,068	$	6,333	$	11,401
Synthetic Biologics, Inc.	293		327		620
Oragenics, Inc.	320		516		836
Fibrocell Science, Inc.	158		61		219
Other	12		618		630
Total	$	5,851	$	7,855	$	13,706

Summary of Deferred Revenue

Deferred revenue consists of the following:

	December 31,
	2014		2013
Upfront and milestone payments	$	107,228	$	72,207
Prepaid research and development services	1,045		1,319
Prepaid product and service revenues	4,365		—
Other	571		45
Total	$	113,209	$	73,571
Current portion of deferred revenue	16,522		7,793
Long-term portion of deferred revenue	96,687		65,778
Total	$	113,209	$	73,571