Exhibit 1

Joint Filing Agreement

In accordance with Rule 13d-1(k) promulgated under the Securities Exchange Act of 1934, as amended, the persons named below agree to the joint filing on behalf of each of them a Statement on Schedule 13D (including amendments thereto) with regard to the common stock of Intrexon Corporation, and further agree that this Joint Filing Agreement be included as an exhibit to such joint filings. In evidence thereof, the undersigned, being duly authorized, hereby execute this agreement as of the 10^th day of June, 2019.

Date:  June 10, 2019

/s/ Randal J. Kirk

Randal J. Kirk

R.J. KIRK DECLARATION OF TRUST
By:	/s/ Randal J. Kirk
	Randal J. Kirk
	Trustee

THIRD SECURITY, LLC
By:	/s/ Randal J. Kirk
	Randal J. Kirk
	Manager

THIRD SECURITY CAPITAL PARTNERS V, LLC
By:	/s/ Randal J. Kirk
	Randal J. Kirk
	Manager

KAPITAL JOE, LLC
By:	/s/ Randal J. Kirk
	Randal J. Kirk
	Manager

NRM VI HOLDINGS I, LLC
By:	/s/ Randal J. Kirk
	Randal J. Kirk
	Manager