X0306 4 2016-12-14 0 0001509190 ARATANA THERAPEUTICS, INC. PETX 0001352447 St Peter Steven C/O ARATANA THERAPEUTICS, INC. 11400 TOMAHAWK CREEK PARKWAY, SUITE 340 LEAWOOD KS 66211 1 1 0 0 PRESIDENT AND CEO Common Stock 2016-12-14 5 G 0 E 50000 0.00 D 592793 D Common Stock 2016-12-15 4 S 0 50000 7.6276 D 542793 D Common Stock 86997 I By LLC Represents charitable gifts of shares by the Reporting Person to two Internal Revenue Code 501(c)(3) non-profit organizations. The sale reported in this Form 4 was effected pursuant to a Rule 10b5-1 trading plan adopted by the Reporting Person. The price reported in Column 4 is a weighted average price. These shares were sold in multiple transactions at prices ranging from $7.60 to $7.73, inclusive. The Reporting Person undertakes to provide to the Issuer, any security holder of the Issuer, or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the ranges set forth in this footnote. Held in the name of Vie Venture LLC, a Delaware limited liability company ("Vie Venture"). The Reporting Person has sole voting and dispositive power over the shares held by Vie Venture, but disclaims beneficial ownership of the shares held thereby except to the extent of his pecuniary interest therein. /s/ Steven St. Peter 2016-12-16