Exhibit 99.1

JOINT FILING AGREEMENT

We, the undersigned, hereby express our agreement that the attached Schedule 13G is filed on behalf of each of us.

Dated: October 10, 2019

THE COLUMN GROUP, LP		THE COLUMN GROUP GP, LP
By:	The Column Group GP, LP
By:	/s/ Peter Svennilson	By:	/s/ Peter Svennilson
Name:	Peter Svennilson	Name:	Peter Svennilson
Title:	Managing Partner	Title:	Managing Partner
PONOI CAPITAL, LP		PONOI MANAGEMENT, LLC
By:	Ponoi Management, LLC
By:	/s/ Peter Svennilson	By:	/s/ Peter Svennilson
Name:	Peter Svennilson	Name:	Peter Svennilson
Title:	Managing Partner	Title:	Managing Partner
PONOI CAPITAL II, LP		PONOI II MANAGEMENT, LLC
By:	Ponoi II Management, LLC
By:	/s/ Peter Svennilson	By:	/s/ Peter Svennilson
Name:	Peter Svennilson	Name:	Peter Svennilson
Title:	Managing Partner	Title:	Managing Partner
By:	/s/ Peter Svennilson
	Peter Svennilson
By:	/s/ David V. Goeddel
	David V. Goeddel
By:	/s/ Tim Kutzkey
	Tim Kutzkey