Collaboration Agreement - Revenue Recognized Related to Each of the Performance Obligations and the Remaining Deferred Revenue (Details) $ in Thousands	6 Months Ended
	Jun. 30, 2022 USD ($)
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Deferred revenue at December 31, 2021	$ 11,715
Milestone	5,000
Revenue recognized	(993)
Deferred revenue at June 30, 2022	15,722
Combined Performance Obligation
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Deferred revenue at December 31, 2021	8,259
Milestone	3,441
Deferred revenue at June 30, 2022	11,700
Development Activities Performance Obligation
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Deferred revenue at December 31, 2021	3,456
Milestone	1,559
Revenue recognized	(993)
Deferred revenue at June 30, 2022	$ 4,022