Collaboration Agreement - Revenue Recognized Related to Each of the Performance Obligations and the Remaining Deferred Revenue (Details) $ in Thousands	3 Months Ended
	Mar. 31, 2022 USD ($)
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Deferred revenue, Beginning balance	$ 11,715
Milestone	468
Revenue recognized	(468)
Deferred revenue, Ending balance	16,247
Combined Performance Obligation
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Deferred revenue, Beginning balance	8,259
Deferred revenue, Ending balance	11,700
Development Activities Performance Obligation
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Deferred revenue, Beginning balance	3,456
Revenue recognized	(468)
Deferred revenue, Ending balance	4,547
Milestone
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Milestone	5,000
Milestone | Combined Performance Obligation
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Milestone	3,441
Milestone | Development Activities Performance Obligation
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items]
Milestone	$ 1,559