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Restructuring Costs (Tables)
 12 Months Ended
Dec. 31, 2016
Restructuring and Related Activities [Abstract]  
Summary of Restructuring Activities

The following table summarizes restructuring activities for the year ended December 31, 2016:

 

     Contract
Termination		      Employee
Termination		      Asset
Impairment		      Total  
     (in thousands)  

Accrual balance at December 31, 2015

    $ —        $ —        $ —        $ —    

Restructuring costs charged to research and development expense

      2,249        —          —          2,249  

Restructuring costs charged to general and administrative expense

      5        332        130        467  

Non-cash charges

      (681 )       —          (130 )       (811 ) 

Cash payments

      (909 )       (233 )       —          (1,142 ) 
  

 

 

     

 

 

     

 

 

     

 

 

  

Accrual balance at December 31, 2016

    $ 664      $ 99      $ —        $ 763