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Restructuring Costs (Tables)
 6 Months Ended
Jun. 30, 2016
Restructuring and Related Activities [Abstract]  
Summary of Restructuring Activities

The following table summarizes restructuring activities for the six months ended June 30, 2016:

 

     Contract
Termination		      Employee
Termination		      Asset
Impairment		      Total  
     (in thousands)  

Accrual balance at December 31, 2015

    $ —         $ —         $ —         $ —     

Restructuring costs charged to research and development expense

      2,787         —           —           2,787   

Restructuring costs charged to general and administrative expense

      5         186         130         321   

Non-cash charges

      (681 )       —           (130 )       (811 ) 

Cash payments

      (18 )       (129 )       —           (147 ) 
  

 

 

     

 

 

     

 

 

     

 

 

  

Accrual balance at June 30, 2016

    $ 2,093       $ 57       $ —         $ 2,150