SEC FORM 3

SEC Form 3

FORM 3

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549

INITIAL STATEMENT OF BENEFICIAL OWNERSHIP OF SECURITIES

Filed pursuant to Section 16(a) of the Securities Exchange Act of 1934
or Section 30(h) of the Investment Company Act of 1940

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1. Name and Address of Reporting Person^*

MARCUS JOEL S

(Last)

(First)

(Middle)

430 E. 29TH STREET, 10TH FLOOR

(Street)

NEW YORK

10016

(City)

(State)

(Zip)

2. Date of Event Requiring Statement (Month/Day/Year)
06/08/2018

3. Issuer Name and Ticker or Trading Symbol
MeiraGTx Holdings plc [ MGTX ]

4. Relationship of Reporting Person(s) to Issuer
(Check all applicable)

X	Director	10% Owner
	Officer (give title below)	Other (specify below)

5. If Amendment, Date of Original Filed (Month/Day/Year)

6. Individual or Joint/Group Filing (Check Applicable Line)

X	Form filed by One Reporting Person
	Form filed by More than One Reporting Person

Table I - Non-Derivative Securities Beneficially Owned
1. Title of Security (Instr. 4)	2. Amount of Securities Beneficially Owned (Instr. 4)	3. Ownership Form: Direct (D) or Indirect (I) (Instr. 5)	4. Nature of Indirect Beneficial Ownership (Instr. 5)
Ordinary Shares	1,649,393	I	See footnote⁽¹⁾
Ordinary Shares	79,922	I	By Third Amended and Restated Joel and Barbara Marcus Family Trust

Table II - Derivative Securities Beneficially Owned (e.g., puts, calls, warrants, options, convertible securities)
1. Title of Derivative Security (Instr. 4)	2. Date Exercisable and Expiration Date (Month/Day/Year)		3. Title and Amount of Securities Underlying Derivative Security (Instr. 4)		4. Conversion or Exercise Price of Derivative Security	5. Ownership Form: Direct (D) or Indirect (I) (Instr. 5)	6. Nature of Indirect Beneficial Ownership (Instr. 5)
1. Title of Derivative Security (Instr. 4)	Date Exercisable	Expiration Date	Title	Amount or Number of Shares	4. Conversion or Exercise Price of Derivative Security	5. Ownership Form: Direct (D) or Indirect (I) (Instr. 5)	6. Nature of Indirect Beneficial Ownership (Instr. 5)
Stock Option	⁽²⁾	03/04/2026	Ordinary Shares	12,883	$7.72	D
Stock Option	⁽³⁾	01/10/2028	Ordinary Shares	15,459	$5.63	D

Explanation of Responses:

1. Represents securities held of record by Alexandria Equities No. 7 ("Alexandria"). ARE-QRS Corp. is the general partner of Alexandria Real Estate Equities L.P., which is the managing member of ARE-Special Services, LLC, which is the managing member of Alexandria. ARE-QRS Corp. has full voting and investment power with respect to the shares owned by Alexandria. The reporting person is the Executive Chairman of ARE-QRS Corp. and may therefore be deemed to have voting and investment power with respect to the shares owned by Alexandria. Each of the reporting person, ARE-QRS Corp., Alexandria Real Estate Equities L.P., and ARE-Special Services, LLC disclaims beneficial ownership of such shares except to the extent of any pecuniary interest therein.

2. The option fully vested with respect to half of the total grant on March 4, 2016 and as to the remainder of the grant on March 4, 2017.

3. The option will vest and become exercisable on January 10, 2019.

Remarks:

Exhibit 24 - Power of Attorney.

	/s/ Richard Giroux, Attorney-in-Fact for Joel S. Marcus	06/08/2018
	** Signature of Reporting Person	Date
Reminder: Report on a separate line for each class of securities beneficially owned directly or indirectly.
* If the form is filed by more than one reporting person, see Instruction 5 (b)(v).
** Intentional misstatements or omissions of facts constitute Federal Criminal Violations See 18 U.S.C. 1001 and 15 U.S.C. 78ff(a).
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