Balance Sheets Details - Property and Equipment, net (Details) - USD ($) $ in Thousands	Jun. 30, 2019	Dec. 31, 2018
Property, Plant and Equipment [Line Items]
Property and equipment	$ 38,579	$ 37,852
Less: accumulated depreciation and amortization	(33,267)	(33,093)
Property and equipment, net	5,312	4,759
Laboratory equipment [Member]
Property, Plant and Equipment [Line Items]
Property and equipment	22,464	21,328
Leasehold improvements [Member]
Property, Plant and Equipment [Line Items]
Property and equipment	10,804	10,359
Computer equipment and software [Member]
Property, Plant and Equipment [Line Items]
Property and equipment	3,772	3,954
Office equipment and furniture [Member]
Property, Plant and Equipment [Line Items]
Property and equipment	1,402	1,272
Construction in progress [Member]
Property, Plant and Equipment [Line Items]
Property and equipment	$ 137	$ 939

Amount of accumulated depreciation, depletion and amortization for physical assets used in the normal conduct of business to produce goods and services.

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Amount before accumulated depreciation, depletion and amortization of physical assets used in the normal conduct of business and not intended for resale. Examples include, but are not limited to, land, buildings, machinery and equipment, office equipment, and furniture and fixtures.

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