X0202 SCHEDULE 13G/A 0000935836-26-000156 0001105806 XXXXXXXX LIVE 1 Common Stock 06/30/2026 0001200375 Codexis, Inc. 192005106 200 Penobscot Drive Redwood City CA 94063 Rule 13d-1(c) LYTTON LAURENCE W X1 5498680.00 94292.00 5498680.00 94292.00 5592972.00 N 5.2 IN The reporting persons are reporting their holdings as of the date of this filing. As of June 30, 2026, the reporting person's beneficial ownership was less than 5%, but subsequent purchases caused his beneficial ownership to increase. Percentage calculated based on 107,561,409 shares of Common Stock outstanding following the issuance of shares by the Issuer described in the Prospectus filed by the Issuer on July 24, 2026. Codexis, Inc. 200 Penobscot Drive, Redwood City, CA 94063 Laurence W. Lytton 467 Central Park West New York, NY 10025 USA Y 5,592,972 5.2% 5,498,680 94,262 5,498,680 94,262 Y Y Y Y Y N By signing below I certify that, to the best of my knowledge and belief, the securities referred to above were not acquired and are not held for the purpose of or with the effect of changing or influencing the control of the issuer of the securities and were not acquired and are not held in connection with or as a participant in any transaction having that purpose or effect, other than activities solely in connection with a nomination under § 240.14a-11. LYTTON LAURENCE W /s/ Laurence W Lytton 08/13/2026