Commitments and Contingencies - Additional Information (Details) - USD ($)	1 Months Ended	9 Months Ended
	Jun. 30, 2012	Sep. 30, 2016	Feb. 29, 2016	Dec. 31, 2015
Commitments And Contingencies [Line Items]
Lease expiration term		The Company leases office and laboratory space under two non-cancelable operating leases in Tucson, Arizona.
Lease extension term, in years		5 years
Expected increase in lease rent		$ 804,000
Incentive from landlord		710,000
Leasehold improvements		1,847,378	$ 710,000	$ 224,061
Monthly accreted lease term as reduction of rent expense		11,833
Capital lease commitments		148,900
Lease equipment financing arrangement		$ 36,000
Lease expiration date of lease payments not in schedule		2017-12
Merck Sharp And Dohme Corporation [Member]
Commitments And Contingencies [Line Items]
Non-exclusive license agreement, milestone accrued or paid	$ 0
Minimum
Commitments And Contingencies [Line Items]
Capital lease commitments of computer equipment period		36 months
Minimum | Merck Sharp And Dohme Corporation [Member]
Commitments And Contingencies [Line Items]
Non-exclusive license agreement, contingent milestone payments	$ 50,000
Non-exclusive license agreement, future royalties	3.00%
Maximum
Commitments And Contingencies [Line Items]
Capital lease commitments of computer equipment period		48 months
Maximum | Merck Sharp And Dohme Corporation [Member]
Commitments And Contingencies [Line Items]
Non-exclusive license agreement, contingent milestone payments	$ 1,000,000
Non-exclusive license agreement, future royalties	6.00%