X0306 4 2015-08-24 0 0001497253 ORGANOVO HOLDINGS, INC. ONVO 0001162525 MICHAELS BARRY D 6275 NANCY RIDGE DRIVE, SUITE 110 SAN DIEGO CA 92121 0 1 0 0 Chief Financial Officer Common Stock 2015-08-24 4 F 0 100692 2.32 D 86808 D Common Stock 285646 I Spouse Stock Option (Right to Buy) 2.32 2015-08-24 4 A 0 100692 0.00 A 2015-08-24 2025-08-24 Common Stock 100692 100692 D The shares were withheld at the election of the Reporting Person to satisfy tax withholding obligations in connection with the partial vesting of restricted stock units granted in August 2012 in accordance with the terms of the Reporting Person's Equity Incentive Award Agreement dated August 2012. This number does not include 1,014,693 shares of common stock underlying outstanding options held by Mr. Michaels; post-transaction Mr. Michaels continues to beneficially own or have a right to acquire 1,387,147 shares of common stock (including the 285,646 shares held by spouse). The option was automatically issued pursuant to the terms of the Reporting Person's Equity Incentive Award Agreement dated August 2012. /s/ Barry Michaels 2015-08-24