NIH Contract (Details) - USD ($) $ in Thousands	Dec. 31, 2021	Dec. 31, 2020
Research and Development [Abstract]
Cumulative cash receipts from milestones achieved	$ 34,016	$ 25,436
Cumulative amounts applied against operating costs (excluding depreciation)	(4,522)	(1,488)
Cumulative amounts applied against depreciation expense for assets placed in service	(703)	0
Cumulative amounts recognized as non-operating income	(7,140)	0
Total deferred grant income	21,651	23,948
Assets placed in service, gross	16,890	0
Construction-in-progress	3,909	9,652
Cumulative amounts applied against depreciation expense for assets placed in service	(703)	0
Carrying value of property and equipment, net	20,096	9,652
Estimated future operating costs, excluding depreciation	0	2,912
Estimated future capital expenditures	1,555	11,384
Deferred grant income, current	3,535	2,912
Deferred grant income, non-current	$ 18,116	$ 21,036