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NIH Contract (Tables)
12 Months Ended
Dec. 31, 2021
Research and Development [Abstract]  
Research and Development Arrangement, Contract to Perform for Others
The following tables summarize the activity under the NIH Contract through December 31, 2021 and 2020 (in thousands):
Year Ended December 31,
20212020
Cumulative cash receipts from milestones achieved$34,016 $25,436 
Cumulative amounts applied against operating costs (excluding depreciation)(4,522)(1,488)
Cumulative amounts applied against depreciation expense for assets placed in service(703)— 
Cumulative amounts recognized as non-operating income(7,140)— 
Total deferred grant income$21,651 $23,948 
Assets placed in service, gross$16,890 $— 
Construction-in-progress3,909 9,652 
Cumulative amounts applied against depreciation expense(703)— 
Carrying value of property and equipment, net20,096 9,652 
Estimated future operating costs, excluding depreciation— 2,912 
Estimated future capital expenditures1,555 11,384 
Total deferred grant income$21,651 $23,948 
Deferred grant income, current$3,535 $2,912 
Deferred grant income, non-current18,116 21,036 
Total deferred grant income$21,651 $23,948