Investments and Fair Value Measurements - Marketable Securities (Details) - USD ($) $ in Thousands	Sep. 30, 2018	Dec. 31, 2017
Cash equivalents:
Cash and cash equivalents	$ 770,036	$ 779,345
Short-term investments:
Short-term investments, amortized cost	254,924	195,992
Short-term investments, gross unrealized gain	50	6
Short-term investments, gross unrealized loss	(1,153)	(464)
Short-term investments, fair value	253,821	195,534
Cash equivalents and short-term investments, carrying amount	280,925	276,875
Gross unrealized gain	50	6
Gross unrealized loss	(1,157)	(467)
Cash equivalents and short-term investments, fair value	279,818	276,414
Level 1
Short-term investments:
Short-term investments, fair value	28,823	17,431
Cash equivalents and short-term investments, fair value	29,877	76,384
Level 2
Short-term investments:
Short-term investments, fair value	224,998	178,103
Cash equivalents and short-term investments, fair value	249,941	200,030
U.S. treasury securities
Short-term investments:
Short-term investments, amortized cost	28,967	17,472
Short-term investments, gross unrealized gain	0	0
Short-term investments, gross unrealized loss	(144)	(41)
Short-term investments, fair value	28,823	17,431
U.S. treasury securities \| Level 1
Short-term investments:
Short-term investments, fair value	28,823	17,431
U.S. treasury securities \| Level 2
Short-term investments:
Short-term investments, fair value	0	0
U.S. government agency securities
Short-term investments:
Short-term investments, amortized cost	14,315	11,540
Short-term investments, gross unrealized gain	0	0
Short-term investments, gross unrealized loss	(85)	(30)
Short-term investments, fair value	14,230	11,510
U.S. government agency securities \| Level 1
Short-term investments:
Short-term investments, fair value	0	0
U.S. government agency securities \| Level 2
Short-term investments:
Short-term investments, fair value	14,230	11,510
Municipal bonds
Short-term investments:
Short-term investments, amortized cost	4,961	6,974
Short-term investments, gross unrealized gain		2
Short-term investments, gross unrealized loss	(14)	(8)
Short-term investments, fair value	4,947	6,968
Municipal bonds \| Level 1
Short-term investments:
Short-term investments, fair value	0	0
Municipal bonds \| Level 2
Short-term investments:
Short-term investments, fair value	4,947	6,968
Foreign government securities
Short-term investments:
Short-term investments, amortized cost	5,020	7,499
Short-term investments, gross unrealized gain	0	0
Short-term investments, gross unrealized loss	(15)	(27)
Short-term investments, fair value	5,005	7,472
Foreign government securities \| Level 1
Short-term investments:
Short-term investments, fair value	0	0
Foreign government securities \| Level 2
Short-term investments:
Short-term investments, fair value	5,005	7,472
Corporate debt securities
Short-term investments:
Short-term investments, amortized cost	145,074	105,144
Short-term investments, gross unrealized gain	47	3
Short-term investments, gross unrealized loss	(684)	(273)
Short-term investments, fair value	144,437	104,874
Corporate debt securities \| Level 1
Short-term investments:
Short-term investments, fair value	0	0
Corporate debt securities \| Level 2
Short-term investments:
Short-term investments, fair value	144,437	104,874
Commercial papers
Short-term investments:
Short-term investments, amortized cost	21,076	30,798
Short-term investments, gross unrealized gain	0	1
Short-term investments, gross unrealized loss	(6)	(9)
Short-term investments, fair value	21,070	30,790
Commercial papers \| Level 1
Short-term investments:
Short-term investments, fair value	0	0
Commercial papers \| Level 2
Short-term investments:
Short-term investments, fair value	21,070	30,790
Certificates of deposit
Short-term investments:
Short-term investments, amortized cost	1,258	1,255
Short-term investments, gross unrealized gain	0	0
Short-term investments, gross unrealized loss	0	0
Short-term investments, fair value	1,258	1,255
Certificates of deposit \| Level 1
Short-term investments:
Short-term investments, fair value	0	0
Certificates of deposit \| Level 2
Short-term investments:
Short-term investments, fair value	1,258	1,255
Asset-backed securities
Short-term investments:
Short-term investments, amortized cost	34,253	15,310
Short-term investments, gross unrealized gain	3	0
Short-term investments, gross unrealized loss	(205)	(76)
Short-term investments, fair value	34,051	15,234
Asset-backed securities \| Level 1
Short-term investments:
Short-term investments, fair value	0	0
Asset-backed securities \| Level 2
Short-term investments:
Short-term investments, fair value	34,051	15,234
Money market funds
Cash equivalents:
Cash and cash equivalents	1,054	58,953
Cash equivalents, fair value	1,054	58,953
Money market funds \| Level 1
Cash equivalents:
Cash equivalents, fair value	1,054	58,953
Money market funds \| Level 2
Cash equivalents:
Cash equivalents, fair value	0	0
Commercial papers
Cash equivalents:
Cash and cash equivalents	22,428	21,930
Cash equivalents, gross unrealized gain	0	0
Cash equivalents, gross unrealized loss	(4)	(3)
Cash equivalents, fair value	22,424	21,927
Commercial papers \| Level 1
Cash equivalents:
Cash equivalents, fair value	0	0
Commercial papers \| Level 2
Cash equivalents:
Cash equivalents, fair value	22,424	21,927
Municipal bonds
Cash equivalents:
Cash and cash equivalents	2,519
Cash equivalents, gross unrealized gain	0
Cash equivalents, gross unrealized loss	0
Cash equivalents, fair value	2,519
Municipal bonds \| Level 1
Cash equivalents:
Cash equivalents, fair value	0
Municipal bonds \| Level 2
Cash equivalents:
Cash equivalents, fair value	2,519
Cash equivalents
Cash equivalents:
Cash and cash equivalents	26,001	80,883
Cash equivalents, fair value	25,997	80,880
Cash equivalents \| Level 1
Cash equivalents:
Cash equivalents, fair value	1,054	58,953
Cash equivalents \| Level 2
Cash equivalents:
Cash equivalents, fair value	$ 24,943	$ 21,927

Fair value portion of probable future economic benefits obtained or controlled by an entity as a result of past transactions or events.

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Amount of investments in debt securities classified as available-for-sale, expected to be converted to cash, sold or exchanged within one year or the normal operating cycle, if longer.

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Cash and Cash Equivalents, Accumulated Gross Unrealized Gain

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