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Stock-Based Compensation - Employee and Director Stock Option Activity (Details) - Employees, Directors and Consultants
3 Months Ended
Mar. 31, 2022
$ / shares
shares
Number of Shares  
Beginning balance (in shares) | shares 3,618,145
Granted (in shares) | shares 332,296
Forfeited (in shares) | shares (444,972)
Ending balance (in shares) | shares 3,505,469
Vested (in shares) | shares 1,089,599
Exercisable (in shares) | shares 1,089,599
Weighted Average Exercise Price  
Beginning balance (in dollars per share) | $ / shares $ 13.55
Granted (in dollars per share) | $ / shares 8.72
Forfeited (in dollars per share) | $ / shares 16.28
Ending balance (in dollars per share) | $ / shares 12.74
Vested (in dollars per share) | $ / shares 14.19
Exercisable (in dollars per share) | $ / shares $ 14.19