Collaboration and Other Agreements - NIAID Contract (Details) $ in Thousands		3 Months Ended		6 Months Ended
	Sep. 15, 2015 Molecule	Jun. 30, 2022 USD ($)	Jun. 30, 2021 USD ($)	Jun. 30, 2022 USD ($)	Jun. 30, 2021 USD ($)
Collaboration And Other Agreements [Line Items]
Revenues		$ 26,007	$ 30,757	$ 37,108	$ 47,638
NIAID
Collaboration And Other Agreements [Line Items]
Number of commercialized molecules | Molecule	2
Total potential value				25,100
Revenues From Grants | NIAID
Collaboration And Other Agreements [Line Items]
Revenues		$ 500	$ 400	$ 900	$ 1,200