X0306 4 2019-10-31 0 0001123494 HARVARD BIOSCIENCE INC HBIO 0001793157 Singh Yash C/O HARVARD BIOSCIENCE, INC. 84 OCTOBER HILL ROAD HOLLISTON MA 01746 0 1 0 0 Executive Vice President Common Stock 2019-10-31 4 A 0 39286 0.00 A 39286 D Stock option (right to buy) 2.80 2019-10-31 4 A 0 86614 0.00 A 2020-10-31 2029-10-31 Common Stock, par value $0.01 per share 86614 86614 D Represents a deferred stock award of 39,286 restricted stock units with performance based vesting conditions. These restricted stock units shall vest in equal installments on October 31, 2020, 2021 and 2022, and are linked to the achievement of a relative total shareholder return of the Issuer's common stock from October 31, 2019 to the earlier of (i) October 31, 2020 or (ii) upon a change of control (measured relative to the NASDAQ Biotechnology index and based on the 20-day trading average price before each such date, or with respect to a change of control, the per share purchase price in the change of control). The target number of these restricted stock units that may be earned is reported above; the maximum amount is 150% of the number reported. Includes the award described in footnote (1) above. The options vest in four equal installments on October 31, 2020, 2021, 2022 and 2023. This form has been signed under power of attorney. /s/ Chad Porter, by power of attorney 2019-11-05