X0306 4 2019-07-07 1 0001123494 HARVARD BIOSCIENCE INC HBIO 0001585223 Duchemin Jeffrey C/O HARVARD BIOSCIENCE, INC. 84 OCTOBER HILL ROAD HOLLISTON MA 01746 1 1 0 0 Chief Executive Officer Common Stock 2019-07-07 4 D 0 285473 0.00 D 464528 D The following restricted stock units were forfeited as a result of the Reporting Person's separation of service from the Issuer effective as of July 7, 2019: (a) 89,402 restricted stock units granted to the Reporting Person on May 25, 2017; (b) 47,222 restricted stock units granted to the Reporting Person on May 24, 2018; (c) 7,556 restricted stock units granted to the Reporting Person on May 24, 2018 that related to an award with a performance condition tied to relative total shareholder return; (d) 84,776 restricted stock units granted to the Reporting Person on March 7, 2019; and (e) restricted stock units, with a target amount of 56,517, granted to the Reporting Person on March 7, 2019 contingent upon achievement of a performance condition tied to relative total shareholder return. This form has been signed under power of attorney. /s/ Chad Porter, by power of attorney 2019-07-09