<?xml version="1.0" encoding="UTF-8"?><edgarSubmission xmlns="http://www.sec.gov/edgar/ownership" xmlns:com="http://www.sec.gov/edgar/common">
  <headerData>
    <submissionType>144</submissionType>
    <filerInfo>
      <filer>
        <filerCredentials>
          <cik>0001695434</cik>
          <ccc>XXXXXXXX</ccc>
        </filerCredentials>
      </filer>
      <liveTestFlag>LIVE</liveTestFlag>



    </filerInfo>
  </headerData>
  <formData>
    <issuerInfo>
      <issuerCik>0001069258</issuerCik>
      <issuerName>KRATOS DEFENSE &amp; SECURITY SOLUTIONS, INC.</issuerName>
      <secFileNumber>001-34460</secFileNumber>
      <issuerAddress>
        <com:street1>10680 TREENA STREET</com:street1>
        <com:street2>SUITE 600</com:street2>
        <com:city>SAN DIEGO</com:city>
        <com:stateOrCountry>CA</com:stateOrCountry>
        <com:zipCode>92131</com:zipCode>
      </issuerAddress>
      <issuerContactPhone>858-812-7300</issuerContactPhone>
      <nameOfPersonForWhoseAccountTheSecuritiesAreToBeSold>Fendley Steven S.</nameOfPersonForWhoseAccountTheSecuritiesAreToBeSold>
      <relationshipsToIssuer>
        <relationshipToIssuer>Officer</relationshipToIssuer>
      </relationshipsToIssuer>
    </issuerInfo>
    <securitiesInformation>
      <securitiesClassTitle>Common Stock</securitiesClassTitle>
      <brokerOrMarketmakerDetails>
        <name>Fidelity Brokerage Services LLC</name>
        <address>
          <com:street1>245 Summer Street</com:street1>
          <com:city>Boston</com:city>
          <com:stateOrCountry>MA</com:stateOrCountry>
          <com:zipCode>02110</com:zipCode>
        </address>
      </brokerOrMarketmakerDetails>
      <noOfUnitsSold>7000</noOfUnitsSold>
      <aggregateMarketValue>487119.93</aggregateMarketValue>
      <noOfUnitsOutstanding>168840708</noOfUnitsOutstanding>
      <approxSaleDate>03/30/2026</approxSaleDate>
      <securitiesExchangeName>NASDAQ</securitiesExchangeName>
    </securitiesInformation>
    <securitiesToBeSold>
      <securitiesClassTitle>Common Stock</securitiesClassTitle>
      <acquiredDate>02/28/2025</acquiredDate>
      <natureOfAcquisitionTransaction>Share Grant</natureOfAcquisitionTransaction>
      <nameOfPersonfromWhomAcquired>Kratos Defense &amp; Security Solutions, Inc.</nameOfPersonfromWhomAcquired>
      <isGiftTransaction>N</isGiftTransaction>
      <amountOfSecuritiesAcquired>7000</amountOfSecuritiesAcquired>
      <paymentDate>02/28/2025</paymentDate>
      <natureOfPayment>N/A</natureOfPayment>
    </securitiesToBeSold>
    <nothingToReportFlagOnSecuritiesSoldInPast3Months>N</nothingToReportFlagOnSecuritiesSoldInPast3Months>
    <securitiesSoldInPast3Months>
      <sellerDetails>
        <name>Steven S. Fendley c/o Kratos Defense &amp; Security Solutions</name>
        <address>
          <com:street1>1 Chisholm Trail,, Suite 300</com:street1>
          <com:city>Round Rock</com:city>
          <com:stateOrCountry>TX</com:stateOrCountry>
          <com:zipCode>78681</com:zipCode>
        </address>
      </sellerDetails>
      <securitiesClassTitle>Common Stock</securitiesClassTitle>
      <saleDate>01/26/2026</saleDate>
      <amountOfSecuritiesSold>7000</amountOfSecuritiesSold>
      <grossProceeds>755824.02</grossProceeds>
    </securitiesSoldInPast3Months>
    <securitiesSoldInPast3Months>
      <sellerDetails>
        <name>Steven S. Fendley c/o Kratos Defense &amp; Security Solutions</name>
        <address>
          <com:street1>1 Chisholm Trail,, Suite 300,</com:street1>
          <com:city>Round Rock</com:city>
          <com:stateOrCountry>TX</com:stateOrCountry>
          <com:zipCode>78681</com:zipCode>
        </address>
      </sellerDetails>
      <securitiesClassTitle>Common Stock</securitiesClassTitle>
      <saleDate>02/23/2026</saleDate>
      <amountOfSecuritiesSold>7000</amountOfSecuritiesSold>
      <grossProceeds>647976.30</grossProceeds>
    </securitiesSoldInPast3Months>
    <securitiesSoldInPast3Months>
      <sellerDetails>
        <name>Steven S. Fendley c/o Kratos Defense &amp; Security Solutions</name>
        <address>
          <com:street1>1 Chisholm Trail,, Suite 300</com:street1>
          <com:city>Round Rock</com:city>
          <com:stateOrCountry>TX</com:stateOrCountry>
          <com:zipCode>78681</com:zipCode>
        </address>
      </sellerDetails>
      <securitiesClassTitle>Common Stock</securitiesClassTitle>
      <saleDate>12/29/2025</saleDate>
      <amountOfSecuritiesSold>7000</amountOfSecuritiesSold>
      <grossProceeds>535962.04</grossProceeds>
    </securitiesSoldInPast3Months>
    <remarks>shares are being sold from THE STEVEN FENDLEY REV TR U/A 11/6/23</remarks>
    <noticeSignature>
      <noticeDate>03/30/2026</noticeDate>
      <planAdoptionDates>
        <planAdoptionDate>05/12/2025</planAdoptionDate>
      </planAdoptionDates>
      <signature>Audrey Skillern, as a duly authorized representative of Fidelity Brokerage Services LLC, as attorney-in-fact for Steven S. Fendley</signature>
    </noticeSignature>
  </formData>
</edgarSubmission>
